| Teaching Interests |  |
| Research with Students | |
| Brown Family Environmental Center | |
| Environmental Studies Concentration | |
| Kokosing Scenic River Association | |
| Professional Societies & links | |
| Ohio Academy of Science | |
| Ohio Biological Survey | |
| Population biology, evolution, animal behavior & environmental studies A.B. '68 Kenyon College Ph.D. '74 Stanford University |
Biology
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